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E-raamat: Physiological Aspects of Legged Terrestrial Locomotion: The Motor and the Machine

  • Formaat: PDF+DRM
  • Ilmumisaeg: 12-Feb-2017
  • Kirjastus: Springer International Publishing AG
  • Keel: eng
  • ISBN-13: 9783319499802
  • Formaat - PDF+DRM
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  • Formaat: PDF+DRM
  • Ilmumisaeg: 12-Feb-2017
  • Kirjastus: Springer International Publishing AG
  • Keel: eng
  • ISBN-13: 9783319499802

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This book offers a succinct but comprehensive description of the mechanics of muscle contraction and legged terrestrial locomotion. It describes on the one hand how the fundamental properties of muscle tissue affect the mechanics of locomotion, and on the other, how the mechanics of locomotion modify the mechanism of muscle operation under different conditions.Further, the book reports on the design and results of experiments conducted with two goals. The first was to describe the physiological function of muscle tissue (which may be considered as the "motor") contracting at a constant length, during shortening, during lengthening, and under a condition that occurs most frequently in the back-and-forth movement of the limbs during locomotion, namely the stretch-shortening cycle of the active muscle. The second objective was to analyze the interaction between the motor and the "machine" (the skeletal lever system) during walking and running in different scenarios with respect to

speed, step frequency, body mass, gravity, age, and pathological gait. The book will be of considerable interest to physiology, biology and physics students, and provides researchers with stimuli for further experimental and analytical work.

Experimental procedures in the study of muscular function.- Functional anatomy of muscle.- Measurements made during or starting from a state of isometric contraction.- Measurements made after stretching the contracting muscle.- Muscle thermodynamics.- External, internal and total mechanical work done to maintain locomotion.- Walking.- Bouncing gaits: running, trotting and hopping.- Effect of speed, step frequency, age and gravity on the mechanics of running.
Part I Muscle: The Motor
1 Experimental Procedures in the Study of Muscle Mechanics
9(16)
1.1 Muscle Chamber and Stimulation
9(1)
1.2 Isometric Contraction
10(1)
1.3 Isotonic Contraction
11(3)
1.4 Isovelocity Contraction
14(1)
1.5 Single Muscle Fiber and Fiber Segment
15(3)
1.6 Response of a System to an Action
18(7)
References
23(2)
2 Functional Anatomy of Muscle
25(10)
2.1 Structures in Series and in Parallel
25(4)
2.2 Localization of the "Motor" and of the Undamped Elastic Elements
29(3)
2.3 Elastic Elements Having the Function of Containing and Centering the Contractile Component
32(3)
References
34(1)
3 Measurements Made During or Starting from a State of Isometric Contraction
35(34)
3.1 Phases of Muscular Contraction Determined on the Whole Muscle
35(1)
3.2 Stress-Strain Diagram of the Apparent Elastic Elements Determined on the Whole Muscle
36(3)
3.3 Twitch, Clonus and Tetanus
39(4)
3.4 Force-Length Relation (Isometric Contraction)
43(3)
3.5 Functional Consequences of the Force-Length Relation
46(3)
3.5.1 Equilibrium Conditions
46(2)
3.5.2 Limitation of the Movement Created by the Sarcomeres
48(1)
3.6 Force-Velocity Relation (Isotonic and Isovelocity Contractions)
49(9)
3.6.1 Experimental Procedure
49(1)
3.6.2 Description of the Force-Velocity Diagram
50(1)
3.6.3 Effect of Muscle Length
51(1)
3.6.4 Force-Velocity of Shortening Relation at Different Times Since the Beginning of Stimulation
52(1)
3.6.5 General Meaning of the Force-Velocity of Shortening Relation
52(2)
3.6.6 Theoretical Interpretation of the Force-Velocity of Shortening Relation
54(4)
3.7 Functional Consequences of the Force-Velocity Relation
58(1)
3.7.1 Power
58(1)
3.7.2 Cost of Positive and Negative Work
59(1)
3.8 Dynamic Force-Length Diagram (Iso-velocity Contraction)
59(2)
3.9 Phases of Muscular Contraction Determined on the Single Muscle Fiber
61(8)
3.9.1 Force-Length Diagram of the Undamped Structure Within the Sarcomere
63(2)
3.9.2 Force-Length Diagram of the Damped Structure Within the Sarcomere
65(1)
References
66(3)
4 Measurements Made After Stretching the Contracting Muscle
69(40)
4.1 Evidence of an Enhancement of Positive Work Production by a Previously Stretched Muscle
69(3)
4.2 What is the Origin of the Extra Work Done by a Previously Stretched Muscle?
72(1)
4.3 Experiments Made on the Whole Muscle
73(13)
4.3.1 Mechanical Work and Efficiency in Isolated Frog and Rat Muscle
73(1)
4.3.2 The Apparent Enhancement of the Contractile Component
74(4)
4.3.3 Modification of the Apparent Elastic Characteristics of Muscle
78(5)
4.3.4 Physiological Meaning of the Modification of the Apparent Elastic Characteristics of Muscle
83(1)
4.3.5 Effect of Temperature on the Kinetics of the Fall in Force After Stretching (Stress-Relaxation)
83(2)
4.3.6 Effect of a Time Interval Between Stretching and Shortening
85(1)
4.4 Experiments Made on the Single Muscular Fiber
86(5)
4.4.1 Effect of Temperature and of the Velocity of Lengthening on the Kinetics of the Fall in Force After Stretching
86(2)
4.4.2 The Four Phases of Shortening Against the Maximal Isometric Force Taking Place After a Ramp Stretch
88(2)
4.4.3 Effect of a Time Interval Between End of Stretching and Release to the Maximal Isometric Force
90(1)
4.5 Experiments Made on a Tendon-Free Segment of the Muscular Fiber
91(12)
4.5.1 Transient Shortening Against the Maximal Isometric Force Is not Due to Stress-Relaxation of Tendons
91(2)
4.5.2 Transient Shortening Against the Maximal Isometric Force Is Independent of the Velocity of Stretching
93(2)
4.5.3 Transient Shortening Against the Maximal Isometric Force Is Independent of Sarcomere Stiffness
95(1)
4.5.4 Transient Shortening Against the Maximal Isometric Force also Occurs When the Ramp Stretch Takes Place on the Ascending Limb of the Force-Length Relation
96(2)
4.5.5 Energy Transfer During Stress Relaxation Following Sarcomere Stretch
98(5)
4.6 Interpretation of the Experimental Results: Conclusive Remarks
103(2)
4.7 Differences Between In Vitro and In Vivo Conditions
105(4)
4.7.1 Characteristics of the Movement Imposed to the Muscle
106(1)
4.7.2 Effect of a Sub Maximal Stimulation
106(1)
References
107(2)
5 Muscle Thermodynamics
109(20)
5.1 Interpretation of the Heat Exchanges Between Muscle and Environment
110(4)
5.2 Methods of Heat Measurement
114(1)
5.3 Resting Heat
115(1)
5.4 Initial Heat
115(6)
5.4.1 Activation and Maintenance Heat
115(2)
5.4.2 Shortening Heat
117(1)
5.4.3 Fenn Effect: A Connection Between Heat Production and the Force-Velocity of Shortening Relation?
117(2)
5.4.4 Heat Production During Forcible Stretching a Contracting Muscle
119(1)
5.4.5 Relaxation Heat
120(1)
5.5 Recovery Heat
121(1)
5.6 Efficiency
122(7)
References
123(6)
Part II Locomotion: Motor--Machine Interaction
6 External, Internal and Total Mechanical Work Done During Locomotion
129(10)
6.1 External Work
129(6)
6.1.1 Mechanical Energy Changes of the Center of Mass During Locomotion
131(3)
6.1.2 Assumptions Made in Calculating External Work from the Force Exerted on the Ground
134(1)
6.2 Internal Work
135(2)
6.3 Total Work
137(2)
References
138(1)
7 Walking
139(28)
7.1 The Pendular Mechanism of Walking: A Way to Reduce External Work
139(1)
7.2 Assessment of the Exchange Between Potential and Kinetic Energy
140(6)
7.3 Phase Shift Between Kinetic and Potential Energy
146(1)
7.4 Within the Step Pendular Energy Transduction in Human Walking
147(1)
7.5 The Mechanism of Walking During Growth
148(2)
7.6 Optimal and Freely Chosen Walking Speed
150(1)
7.7 The Mechanism of Walking in Different Animal Species
151(1)
7.8 Effect of Step Frequency on the Mechanical Power Output in Human Walking
151(5)
7.9 Role of Gravity in Human Walking
156(2)
7.10 Mechanics of Competition Walking
158(3)
7.11 Ergometric Evaluation of Pathological Gait
161(6)
References
164(3)
8 Bouncing Gaits: Running, Trotting and Hopping
167(36)
8.1 Transition from the Mechanism of Walking to the Mechanism of Running
167(1)
8.2 The Bouncing Mechanism of Progression
168(5)
8.3 Vertical Oscillation of the Center of Mass
173(2)
8.4 Physical Division of the Vertical Oscillation of the Center of Mass
175(2)
8.5 The On-Off-Ground Symmetry and Asymmetry of the Rebound
177(3)
8.6 The Landing-Takeoff Symmetry and Asymmetry of the Rebound
180(6)
8.6.1 Height of the Center of Mass at Touch Down and Takeoff
181(1)
8.6.2 The Four Phases of the Bounce and the Transduction of Mechanical Energy During the Running Step
182(3)
8.6.3 Positive and Negative Work Durations
185(1)
8.7 Landing-Takeoff Asymmetry of the Bouncing Step: Asymmetric Motor or Asymmetric Machine?
186(17)
8.7.1 Different Machines with the Same Motor
187(4)
8.7.2 Running Backwards: Soft Landing---Hard Takeoff
191(9)
References
200(3)
9 Effect of Speed, Step Frequency and Age on the Bouncing Step
203(20)
9.1 Effect of the Running Speed on the On-Off-Ground Asymmetry and the Landing-Takeoff Asymmetry
203(3)
9.2 Effect of the Step Frequency on the Mechanical Power Output in Human Running
206(4)
9.3 The Resonant Step Frequency in Human Running
210(2)
9.4 Effect of Age
212(11)
9.4.1 On-Off-Ground Symmetry and Asymmetry
212(1)
9.4.2 Landing-Takeoff Asymmetry During Growth
212(4)
9.4.3 Landing-Takeoff Asymmetry in Old Age
216(4)
References
220(3)
10 Work, Efficiency and Elastic Recovery
223
10.1 Mechanical Work Done by Adult Humans as a Function of Speed
223(7)
10.1.1 External Work
223(2)
10.1.2 Internal Work
225(2)
10.1.3 Total Work and Efficiency
227(3)
10.2 Running Children
230(9)
10.2.1 External Work
230(7)
10.2.2 Internal Work, Total Work and Efficiency
237(2)
10.3 Old Men Running
239(5)
10.4 Effect of Body Mass on Elastic Recovery
244(9)
10.5 Effect of Gravity in Human Running
253(4)
10.6 Sprint Running
257(7)
10.7 Vertical Jump
264
10.7.1 Effect of Stretch Amplitude
266(2)
10.7.2 Jumping at Different Simulated Gravities
268(3)
10.7.3 Metabolic Energy Expenditure
271(1)
References
271
Giovanni A. Cavagna is Emeritus Professor of Human Physiology at State University of Milan, Italy. His research has mainly focused on biomechanics and biophysics.